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Wolfire Games is a tiny studio. Wikipedia says 4 employees. Development of Overgrowth is mostly one person, who documented the whole process in fascinating weekly videos:
https://m.youtube.com/c/WolfireGames/videos
The title says "game engine" because the artwork remains proprietary (similar to old id games, buy the game to get it).
However, just to clarify, all the game code (including gameplay logic, which is sometimes not considered part of the "game engine") is Apache 2.0.
GitHub page: https://github.com/WolfireGames/overgrowth
NOTE:
The number of mentions on this list indicates mentions on common posts plus user suggested alternatives.
Hence, a higher number means a more popular project.