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Apollo released the source code. I think in his case, the developer felt that the project was going to die anyway, and he wanted to clear his name from Reddit's smear campaign.
The main 2 ones for discussion like Reddit are Lemmy and Kbin (fedia.io). And Mastodon that is set up more like Twitter, also works on the same protocol.
NOTE:
The number of mentions on this list indicates mentions on common posts plus user suggested alternatives.
Hence, a higher number means a more popular project.