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I don't know 'all of the underlying technology' but as a little 'maintenance exercise' under the top line file there is an option, "Compact Folders" - you can do them individually but I do it first I clean out my 'trash', then I compact. Although I am on Linux and there is no 'defrag', it probably would not help to compact them. Also, FYI, https://www.bleachbit.org/ program for Windows, Linux, etc. has an option to VACUUM (essentially to tighten up the SQLITE files - databases, I think.) If you are doing Thunderbird I would not let it do anything but the vacuum option (note Thunderbird uses an Mbox file system) but I think the option is still called vacuum in Bleachbit for simplicity sake) until you knew what it was doing, but Firefox also uses SQLITE files and you can vacuum those - for that program I let it wipe everything and just select every box depending if I am running as root-superuser-admin or not.) If you just converted from one .... perhaps everything is spread across your drive. NOTE: if your drive is a SSD you are NOT supposed to have to, nor is it suggested that you defrag (or do similar actions) to your drive to preserve the life of the SSD. In which case you might want to understand the difference between 'compact' and defrag. Please do appropriate research on all issues.
https://support.mozilla.org/en-US/kb/compacting-folders https://www.crucial.com/articles/about-ssd/should-you-defrag-an-ssd Kind of interesting to poke around Firefox's and other programs sqlite database files, just be cautious. https://sqlitebrowser.org/